Craig Woodard, Ph.D.

Professor of Biological Sciences


B.S. in Biology, Bates College, Lewiston , ME 1986
M.S. in Biology, Yale University, December, 1988
M.Phil. in Biology, Yale University, December, 1988
Ph.D. in Biology, Yale University, November, 1992

Postdoctoral Training

Laboratory of Dr. Carl S. Thummel, Howard Hughes Medical Institute and Department of Human Genetics, University of Utah, September, 1992 - August, 1995

Research Interests

  1. Control of animal development by steroid hormones
  2. Genetic regulation of sexual behavior


  • Biology 145  Biology in the Age of the Human Genome Project  (spring)
  • Biology 210  Genetics and Molecular Biology  (fall)
  • Biology 340  Eukaryotic Molecular Biology  (spring)

Selected Publications

Packard, B. W., Marciano, V., Payne, J.M., Bledzki, L. A., Woodard, C.T.
(2014). Negotiating Peer Mentoring Roles in Undergraduate Research Lab Settings. Mentoring & Tutoring: Partnership in Learning. doi: 10.1080/13611267.2014.983327

Bond, N., Nelliot, A., Bernardo, M.K., Gorski, K., Ayerh, M., Hoshizaki, D.K., and Woodard, C.T. (2011). BFTZ-F1 and Matrix Metalloproteinase 2 are Required for Fat-Body Remodeling in Drosophila.  Developmental Biology. doi:10.1016/j.ydbio.2011.09.015

Kim, C., Srivastava, S., Rice, M., Godenschwege, T.A., Bentley, B., Shao. S., Ravi, S., Woodard, C.T., and Schwartz, L.M. (2011). Expression of human amyloid precursor protein in the skeletal muscles of Drosphila results in age- and activity-dependent muscle weakness. BMC Physiology. 11:7. doi:10.1186/1472-6793-11-7.

Maloney, M., Parker, J., LeBlanc, M., Woodard, C.T., Glackin, M., and Hanrahan, M. (2010). Bioinformatics and the Undergraduate Curriculum. CBE-Life Sciences Education 9:172-174.

Woodard, C., Alcorta, E., and Carlson, J. (2007). The rdgB Gene of  Drosophila: A Link Between Vision and Olfaction, J. Neurogenetics 21:  291-305 (Reissued).

Pick, L., Anderson, W.R., Shulz, J., and Woodard, C.T. 2006. The Ftz-F1 family: orphan nuclear receptors regulated by novel protein-protein interactions. In: Taneja, R., editor. Nuclear receptors in development. Vol. 16. (Advances in Developmental Biology Series). San Diego, CA: Elsevier BV. pp. 255-296.

Fortier, T.M., Chatterjee, R., Klinedinst, S., Baehrecke, E.H., and Woodard, C.T. 2006. how functions in leg development during Drosophila metamorphosis. Developmental Dynamics. 235: 2248-2259.

Fortier, T.M., Vasa, P.P., and Woodard, C.T. 2003. Orphan Nuclear Receptor FTZ-F1 is Required for Muscle-Driven Morphogenetic Events at the Prepupal-Pupal Transition in Drosophila melanogaster. Developmental Biology. 257: 153-165.

Lee C.-Y., Simon C.R., Woodard, C.T. and Baehrecke E.H. 2002. Genetic Mechanism for the Stage- and Tissue-Specific Regulation of Steroid-Triggered Programmed Cell Death in Drosophila. Developmental Biology. 252: 138-148.

Gaines, P, Tompkins, L., Woodard, C.T., and Carlson, J.R. 2000. quick-to-court, a Drosophila Mutant with Elevated Levels of Sexual Behavior, is Defective in a Predicted Coiled-Coil Protein. Genetics 154: 1627-1637.

Gaines, P., Woodard, C.T., and Carlson, J.R. 1999. An Enhancer Trap Line Identifies the Drosophila Homolog of the L37a Ribosomal Protein. Gene 239: 137-143.

Broadus, J., **McCabe, J., Endrizzi, B., Thummel, C.S., and Woodard, C.T. 1999. The Drosophila ßFTZ-F1 orphan nuclear receptor provides competence for stage-specific responses to the steroid hormone ecdysone. Molecular Cell 3: 143-149.

Woodard, C.T., Baehrecke, E.H., and Thummel, C.S. 1994. A Molecular Mechanism for the Stage-Specificity of the Drosophila Prepupal Genetic Response to Ecdysone. Cell 79: 607-615.

Riesgo-Escovar, J., Woodard. C., and Carlson. J. 1994. Olfactory Physiology in the Drosophila Maxillary Palps Requires the Visual System Gene rdgB. J. Comp. Physiol. 175: 687-693.

Baehrecke, E.H., Woodard, C.T., and Thummel, C.S. 1994. Temporal Differences in Ecdysone-Regulated Gene Expression at the Onset of Drosophila Metamorphosis. In Perspectives in Comparative Endocrinology. ed. K.G. Davey, R.E. Peter, and S.S. Tobe) pp. 395-399. National Research Council of Canada, Ottawa.